The Principal LLM Application Engineer will develop and optimize Large Language Models, lead projects, evaluate performance, and collaborate across teams to enhance AI capabilities in cybersecurity.
About System Two Security
System Two Security is driving SOC transformation with its unique application of AI computing with an initial focus on generative AI powered proactive threat research, threat analysis and iterative threat hunting. The product’s purpose-built language agents respond to new threat actors and attack patterns within minutes with its agentic workflows delivering end-to-end threat detection and containment from integrated feeds of raw cyber threat advisories. Early users of the product include prominent MSSPs, and enterprise SOCs across retail, healthcare, SaaS and technology verticals. It is based in Palo Alto, CA and is venture funded by The Hive.
Job Description
System Two Security is looking to hire a Principal LLM Application Engineer where you will focus on leveraging and optimizing Large Language Models (LLMs) along with the implementation of advanced AI technologies. You will be working on cutting edge projects, and will utilize skills in GenAI, LLM and ML Ops to bring a better experience to customers. You will have the opportunity to be a technical leader and to take ownership for projects. It offers a significant opportunity for professional growth at the forefront of AI in the dynamic field of cybersecurity.
Responsibilities
- Development and Optimization of LLMs: Implement and fine-tune state-of-the-art Large Language Models for various applications, focusing on performance and accuracy.
- Evaluating Model Performance: Conduct rigorous evaluations of LLMs, assessing effectiveness, efficiency, and business alignment.
- Integration of Advanced AI Technologies: Implement Retrieval-Augmented Generation (RAG), function calling, and code interpreter technologies to enhance the capabilities of Large Language Models.
- Research and Development: Stay abreast of the latest advancements in machine learning, particularly in LLMs, LLM agents, and large-scale neural network training.
- Data and Model Parallel Training: Utilize data and model parallel training techniques for efficient handling of large-scale models.
- Cross-Functional Collaboration and Leadership: Work with ML engineers, data scientists, and product teams, providing guidance and mentorship.
- Documentation and Reporting: Maintain detailed documentation of methodologies, models, and results, communicating findings across the organization.
- Contribute to product roadmap and vision
- Implement and evaluate various LLM application logic ( flows ) and prompting strategies and stay up to date with the latest advancements in this field
- Lead the incubation of new initiatives, architect scalable solutions, and drive strategic technology choices to develop and deliver AI/ML capabilities in a micoservcies architecture for our customers.
- Design, test, and deploy Machine learning models, including large-language models and build pipelines at scale for batch and real-time use cases.
Required Skills
- Bachelor's degree in Computer Science, Engineering, or related field.
- 5+ years of experience in natural language processing, machine learning, and/or data science.
- Experience in Python or R.
- Experience working with large language models, such as GPT-3+, LLAMA, or similar.
- Strong problem-solving skills and the ability to think creatively to identify new opportunities for LLMs in our products and services.
- Experience with one or more deep learning frameworks.
- A deep theoretical or empirical understanding of deep learning.
- Experience in building, testing and deploying machine learning models, including large language models.
- Strong analytical and debugging skills.
- Familiarity with cloud-based infrastructure and distributed computing.
- Experience with DevOps/MLOps/LLMOps.
- Ability to address complex challenges in model training and optimization.
- Effective communication skills for conveying technical concepts and collaborating with cross-functional teams.
- Passion for staying updated with the latest trends in AI and machine learning.
Preferred Skills
- Master's or PhD in Computer Science, AI, or related fields, with a focus on machine learning and natural language processing.
- Proficiency in programming languages such as Python or R.
- Working experience within Cyber Security.
- Experience in building GenAI solutions using RAG framework and building LLM Agentic applications is preferred.
Benefits
- Medical, Dental, and Vision
- Equity in the company
System Two Security Framingham, Massachusetts, USA Office
Framingham, MA, United States
Similar Jobs
eCommerce • Information Technology • Internet of Things • Mobile • On-Demand • Retail • Software
Own and grow a portfolio of enterprise and upper mid-market clients by driving revenue, retention, and expansion. Build senior stakeholder relationships, execute account strategies, identify cloud and modernization opportunities, manage sales cycles through close, and coordinate solution engineering and delivery teams. Monitor account health, maintain pipeline and forecast accuracy, and ensure successful customer outcomes across managed and professional services engagements.
Top Skills:
AWSAzureGCPVMware
eCommerce • Information Technology • Internet of Things • Mobile • On-Demand • Retail • Software
Manage a portfolio of B2B technology customers, guiding onboarding, adoption, engagement, retention, and renewal readiness. Develop success plans, monitor customer health, conduct business reviews, coordinate cross-functional resources, resolve issues, identify growth opportunities, and maintain customer documentation. The role requires strong relationship management, organization, communication, and collaboration across Sales, Delivery, Operations, Consulting, Engineering, and Support.
Top Skills:
AWSAzureCRMCustomer Success PlatformsGCPVMware
eCommerce • Information Technology • Internet of Things • Mobile • On-Demand • Retail • Software
Owns and grows a portfolio of enterprise and upper mid-market clients by driving revenue, retention, and expansion. Builds senior stakeholder relationships, develops account strategies, manages opportunities through the full sales cycle, and aligns cloud, security, managed services, and professional services solutions to client needs. Coordinates solution engineering and delivery teams, monitors account health, maintains pipeline and forecast accuracy, and supports successful customer outcomes.
Top Skills:
AWSGCPAzureVMware
What you need to know about the Boston Tech Scene
Boston is a powerhouse for technology innovation thanks to world-class research universities like MIT and Harvard and a robust pipeline of venture capital investment. Host to the first telephone call and one of the first general-purpose computers ever put into use, Boston is now a hub for biotechnology, robotics and artificial intelligence — though it’s also home to several B2B software giants. So it’s no surprise that the city consistently ranks among the greatest startup ecosystems in the world.
Key Facts About Boston Tech
- Number of Tech Workers: 269,000; 9.4% of overall workforce (2024 CompTIA survey)
- Major Tech Employers: Thermo Fisher Scientific, Toast, Klaviyo, HubSpot, DraftKings
- Key Industries: Artificial intelligence, biotechnology, robotics, software, aerospace
- Funding Landscape: $15.7 billion in venture capital funding in 2024 (Pitchbook)
- Notable Investors: Summit Partners, Volition Capital, Bain Capital Ventures, MassVentures, Highland Capital Partners
- Research Centers and Universities: MIT, Harvard University, Boston College, Tufts University, Boston University, Northeastern University, Smithsonian Astrophysical Observatory, National Bureau of Economic Research, Broad Institute, Lowell Center for Space Science & Technology, National Emerging Infectious Diseases Laboratories

